I opened up a tac case with cisco on it the rep stated that the 1720 can't do intervlan rotuing bascially it does not support isl or 802.1q. I ordered a 2612 off of ebay last night that will be able ...
I'm not immediately aware of any sort of limitation in the 4000 switch since all you should need to do to set up the inter-VLAN routing with the 5500 switch is establish a VLAN trunk (ISL or 802.1Q) ...
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